Why is swimming more tiring than running?

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Did about 700m worth of swimming in 20 mins today, considerably less time or distance than I normally cover in my 5 or 10k runs, and now exhausted. Admittedly a large part of the excursion involved bundling up Emily, persuading her that pools are fun (she was mostly unconvinced, although did like her snazzy bathing suit), and getting her warm and dry as quickly as possible!

What is it about swimming? Less used muscles, apparently…  but who knows?